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   » » Wiki: Anthracosauria
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Anthracosauria is a order of extinct reptile-like (in the broad sense) that flourished during the and periods, although precisely which species are included depends on one's definition of the taxon. "Anthracosauria" is sometimes used to refer to all more closely related to such as , , and , than to such as and . An equivalent term to this definition would be . Anthracosauria has also been used to refer to a smaller group of large, crocodilian-like aquatic tetrapods also known as .


Various definitions
As originally defined by Säve-Söderbergh in 1934, the anthracosaurs are a group of usually large aquatic Amphibia from the Carboniferous and lower Permian. As defined by Alfred Sherwood Romer however, the anthracosaurs include all non-amniote "" , and Säve-Söderbergh's definition is more equivalent to Romer's suborder . This definition was also used by Edwin H. Colbert and Robert L. Carroll in their textbooks of Vertebrate Palaeontology (Colbert 1969, Carroll 1988). Dr A. L. Panchen however preferred Säve-Söderbergh's original definition of Antracosauria in his Handbuch der Paläoherpetologie, 1970.

With things have changed again. , and Rowe (1988) defined Anthracosauria as a including "Amniota plus all other tetrapods that are more closely related to amniotes than they are to amphibians" (Amphibia in turn was defined by these authors as a clade including and those tetrapods that are more closely related to lissamphibians than they are to ). Similarly, (1996) uses the term in a sense to refer to only the most advanced . Thus his definition includes , Solenodonsauridae and the amniotes. As Ruta, Coates and Quicke (2003) pointed out, this definition is problematic, because, depending on the exact phylogenetic position of Lissamphibia within Tetrapoda, using it might lead to the situation where some taxa traditionally classified as anthracosaurs, including even the genus itself, wouldn't belong to Anthracosauria. Laurin (2001) created a different phylogenetic definition of Anthracosauria, defining it as "the largest clade that includes Anthracosaurus russelli but not ". However, Michael Benton (2000, 2004) makes the anthracosaurs a order within the superorder , along with the orders and , thus making the Anthracosaurians the "lower" reptile-like amphibians. In his definition, the group encompass the , and possibly the family .

Many studies since have suggested that anthracosaurs or embolomeres are likely closer to , but some recent studies either retain them as amphibians or argue that their relationships are still ambiguous and are more likely to be stem-tetrapods.


Etymology
The name "Anthracosauria" is ('coal lizards'), because many of its fossils were found in the .


References and external links


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