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The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of that includes eight extant species in four Genus: Orangutan (the Bornean, Sumatran and Tapanuli orangutan); Gorilla (the Eastern gorilla and western gorilla); Pan (the chimpanzee and the bonobo); and Homo, of which only Human remain.
Numerous revisions in classifying the great apes have caused the use of the term hominid to change over time. The original meaning of "hominid" referred only to humans ( Homo) and their closest extinct relatives. However, by the 1990s humans and other apes were considered to be "hominids".
The earlier restrictive meaning has now been largely assumed by the term Hominini, which comprises all members of the human clade after the split from the chimpanzees ( Pan). The current meaning of "hominid" includes all the great apes including humans. Usage still varies, however, and some scientists and laypersons still use "hominid" in the original restrictive sense; the scholarly literature generally shows the traditional usage until the turn of the 21st century.
Within the taxon Hominidae, a number of extant and extinct genera are grouped with the humans, chimpanzees, and gorillas in the subfamily Homininae; others with orangutans in the subfamily Ponginae (see classification graphic below). The most recent common ancestor of all Hominidae lived roughly 14 million years ago, when the ancestors of the orangutans speciated from the ancestral line of the other three genera.Dawkins R (2004) The Ancestor's Tale. Those ancestors of the family Hominidae had already speciated from the family Hylobatidae (the ), perhaps 15 to 20 million years ago.
Due to the close genetic relationship between humans and the other great apes, certain animal rights organizations, such as the Great Ape Project, argue that nonhuman great apes are persons and should be given basic human rights. Twenty-nine countries have instituted research bans to protect great apes from any kind of scientific testing.
At sites far distant from East Africa, the presence of other generalized non-cercopithecids, that is, non-monkey primates, of middle Miocene age— Otavipithecus from cave deposits in Namibia, and Pierolapithecus and Dryopithecus from France, Spain and Austria—is further evidence of a wide diversity of ancestral ape forms across Africa and the Mediterranean basin during the relatively warm and equable climatic regimes of the early and middle Miocene. The most recent of these far-flung Miocene apes (hominoids) is Oreopithecus, from the fossil-rich coal beds in northern Italy and dated to 9 million years ago.
Molecular evidence indicates that the lineage of gibbons (family Hylobatidae), the "lesser apes", diverged from that of the great apes some 18–12 million years ago, and that of orangutans (subfamily Ponginae) diverged from the other great apes at about 12 million years. There are no fossils that clearly document the ancestry of gibbons, which may have originated in a still-unknown South East Asian hominoid population; but fossil proto-orangutans, dated to around 10 million years ago, may be represented by Sivapithecus from India and Griphopithecus from Turkey. Species close to the last common ancestor of gorillas, chimpanzees and humans may be represented by Nakalipithecus fossils found in Kenya and Ouranopithecus fossils found in Greece. Molecular evidence suggests that between 8 and 4 million years ago, first the gorillas (genus Gorilla), and then the chimpanzees (genus Pan) split off from the line leading to humans. Human DNA is approximately 98.4% identical to that of chimpanzees when comparing single nucleotide polymorphisms (see human evolutionary genetics). The fossil record, however, of gorillas and chimpanzees is limited; both poor preservation—rain forest soils tend to be acidic and dissolve bone—and sampling bias probably contribute most to this problem.
Other hominins probably adapted to the drier environments outside the African equatorial belt; and there they encountered antelope, hyenas, elephants and other forms becoming adapted to surviving in the East African savannas, particularly the regions of the Sahel and the Serengeti. The wet equatorial belt contracted after about 8 million years ago, and there is very little fossil evidence for the divergence of the hominin lineage from that of gorillas and chimpanzees—which split was thought to have occurred around that time. The earliest fossils argued by some to belong to the human lineage are Sahelanthropus tchadensis (7 Ma) and Orrorin tugenensis (6 Ma), followed by Ardipithecus (5.5–4.4 Ma), with species Ar. kadabba and Ar. ramidus.
A number of very similar words apply to related classifications:
A cladogram indicating common names (cf. more detailed cladogram below):
Humans and close relatives including the tribes Hominini and Gorillini form the subfamily Homininae (see classification graphic below). (A few researchers go so far as to refer the chimpanzees and the gorillas to the genus Homo along with humans.) But, those fossil relatives more closely related to humans than the chimpanzees represent the especially close members of the human family, (1986). 9780471626442, Wiley-Liss. ISBN 9780471626442
Many extinct hominids have been studied to help understand the relationship between modern humans and the other extant hominids. Some of the extinct members of this family include Gigantopithecus, Orrorin, Ardipithecus, Kenyanthropus, and the australopithecines Australopithecus and Paranthropus.
The exact criteria for membership in the tribe Hominini under the current understanding of human origins are not clear, but the taxon generally includes those species that share more than 97% of their DNA with the modern human genome, and exhibit a capacity for language or for simple beyond their 'local family' or band. The theory of mind concept—including such faculties as empathy, attribution of mental state, and even empathetic deception—is a controversial criterion; it distinguishes the adult human alone among the hominids. Humans acquire this capacity after about four years of age, whereas it has not been proven (nor has it been disproven) that gorillas or chimpanzees ever develop a theory of mind. This is also the case for some New World monkeys outside the family of great apes, as, for example, the capuchin monkeys.
However, even without the ability to test whether early members of the Hominini (such as Homo erectus, Neanderthal, or even the australopithecines) had a theory of mind, it is difficult to ignore similarities seen in their living cousins. Orangutans have shown the development of culture comparable to that of chimpanzees, and some say the orangutan may also satisfy those criteria for the theory of mind concept. These scientific debates take on political significance for advocates of great ape personhood.
All species are omnivore, although chimpanzees and orangutans primarily eat fruit. When gorillas run short of fruit at certain times of the year or in certain regions, they resort to eating shoots and leaves, often of bamboo, a type of grass. Gorillas have extreme adaptations for chewing and digesting such low-quality forage, but they still prefer fruit when it is available, often going miles out of their way to find especially preferred fruits. Humans, since the Neolithic Revolution, have consumed mostly and other foods, including increasingly highly Food processing, as well as many other domesticated plants (including fruits) and meat.
Both chimpanzees and humans are known to wage over territories and resources.
Gestation in great apes lasts 8–9 months, and results in the birth of a single offspring, or, rarely, twins. The young are born helpless, and require care for long periods of time. Compared with most other mammals, great apes have a remarkably long adolescence, not being weaning for several years, and not becoming fully mature for eight to thirteen years in most species (longer in orangutans and humans). As a result, females typically give birth only once every few years. There is no distinct breeding season.
Gorillas and chimpanzees live in family groups of around five to ten individuals, although much larger groups are sometimes noted. Chimpanzees live in larger groups that break up into smaller groups when fruit becomes less available. When small groups of female chimpanzees go off in separate directions to forage for fruit, the dominant males can no longer control them and the females often mate with other subordinate males. In contrast, groups of gorillas stay together regardless of the availability of fruit. When fruit is hard to find, they resort to eating leaves and shoots.
This fact is related to gorillas' greater sexual dimorphism relative to that of chimpanzees; that is, the difference in size between male and female gorillas is much greater than that between male and female chimpanzees. This enables gorilla males to physically dominate female gorillas more easily. In both chimpanzees and gorillas, the groups include at least one dominant male, and young males leave the group at maturity.
On 25 June 2008, the Spanish parliament supported a new law that would make "keeping apes for circuses, television commercials or filming" illegal. On 8 September 2010, the European Union banned the testing of great apes.
| Bornean orangutan | 104,700 | Critically endangered | |
| Sumatran orangutan | 6,667 | Critically endangered | An estimate of the number of wild orangutans in 2004: |
| Tapanuli orangutan | 800 | Critically endangered | |
| Western gorilla | 200,000 | Critically endangered | |
| Eastern gorilla | <5,000 | Critically endangered | |
| Chimpanzee | 200,000 | Endangered | |
| Bonobo | 10,000 | Endangered | |
| Human | 8,211,817,000 | N/A |
Fig 2: "Divergence time estimates for the jacchus marmoset group based on the BEAST4 (Di Fiore et al., 2015) calibration scheme for alignment A.... Numbers at each node indicate the median divergence time estimate."
Family Hominidae
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